I will sum up my thoughts but Iam not going to get into a lot,
Coyote are first and foremost survivor's,
Why are there populations now moving so rapidly here , pressure they are being forces to move on do to human activity,
Coyote Do not run in packs,,,,,, they run in family units and this unit will brake up at some point in the year and some coyotes will move off to form there own family groups. because of homes and people moving in to more and more areas around are state .it just making the coyote move grater distances to fined there own home Territories, to make it short are communities are growing faster and faster witch is in my opinion why are coyote population is growing so fast here in PA. they are forced to move on to new area.
the more they disperse the grater there numbers grow!
I also believe that it is inherently in he genes for them to survive, to make it short ,
every thing that go's on around them in there environment has some effect on there breeding ,, if coyote numbers are low the food sources are better, so the heather the female the more pups she will have.
I will go as far as to say, I don't think all females in the family unit get bred each year, and if they need to keep there unit nubers up,, do to low population. they will breed more females .Females that might not have had a litter this year will.
hope this makes some scents,,,
and this is just how i look at it's not facts just my opinion